Four Turner men were charged with multiple illegal hunting offenses Monday, following a four-month investigation by the Maine Warden Service, the U.S. Fish and Wildlife Service and the Pennsylvania Game Commission.

The charges followed the search, under warrant, of five homes in Androscoggin and Kennebec counties, according to a statement from Maine Department of Inland Fisheries and Wildlife spokeswoman Deborah Turcotte.

Everett Leonard, 31, was arrested and charged with four violations. His father, E.H. “Lenny” Leonard, 59, was arrested and charged with six violations. Both were booked at the Androscoggin County Jail on Monday but were released on bail later that night, jail officials said. Carlton “John” Enos, 19, and Jason Clifford, 27, were summoned to appear in court on 11 violations each.

The four suspects were linked to the killing of eight deer in Maine and at least 30 in Pennsylvania, Turcotte said. The Warden Service believes the hunters are responsible for nearly 150 violations of Maine’s game laws.

Everett Leonard was charged with night hunting, illuminating wildlife, shooting a firearm from a motor vehicle, and hunting deer after killing one.

E.H. Leonard was charged with two counts of unlawful trafficking in prescription drugs, two counts of criminal trespassing and two counts of unlawfully driving deer.

Carlton was charged with hunting deer in a closed season, possessing an antlerless deer, false registration of deer, trapping without a license, two counts of hunting deer after killing one, two counts of criminal trespassing, two counts of unlawfully driving deer and illegal transportation of deer.

Clifford was charged with two counts of hunting antlerless deer and hunting deer after killing one, possession of antlerless deer and an unregistered deer, two counts of criminal trespassing, two counts of hunting without proper orange and two counts of unlawfully driving deer.

“These individuals are intentional violators who are having a significant impact on Maine and Pennsylvania’s wildlife resources,” Maine Warden Dan Scott said. “They demonstrated complete disregard for wildlife laws, as well as the safety of the public.”

The quartet killed deer for personal use, Turcotte said. “There’s no evidence of commercial use,” she said. “They believed the resources of Maine were theirs for the taking, and obviously they are not.”

During the searches on Monday, game wardens confiscated deer meat, antlers, firearms and bows, ammunition, several taxidermied hawks and owls and a computer, among other items.

The Maine Warden Service initiated the investigation during the summer of 2010 after receiving anonymous tips via the state’s Operation Game Thief hot line that the Leonards were poaching, Scott said.

The Warden Service later began working with the Pennsylvania Game Commission, which alleges that the suspects are behind “hundreds of violations” of hunting laws in Pennsylvania, according to the statement. The Pennsylvania violations occurred mainly in October and December 2010 in Armenia Township, a rural area in northeast Pennsylvania with a population of fewer than 200 people. Wardens believe that the hunters killed deer and brought them back to Maine, Turcotte said.

“We anticipate the suspects will be extradited to Pennsylvania where they will be subject to stiff game laws recently enacted to address just this type of poaching,” Pennsylvania Game Commission Special Operations Chief Thomas Grohol said.